American Philosophical Society
Member History

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2. Biological Sciences[X]
1Name:  Dr. Edgar Douglas Adrian
 Year Elected:  1938
 Class:  2. Biological Sciences
 Residency:  International
 Living? :   Deceased
 Birth Date:  1889
 Death Date:  8/4/77
   
2Name:  Lord Richard Adrian
 Institution:  Cambridge & House of Lords
 Year Elected:  1987
 Class:  2. Biological Sciences
 Subdivision:  201. Molecular Biology and Biochemistry
 Residency:  International
 Living? :   Deceased
 Birth Date:  1927
 Death Date:  4/4/95
   
3Name:  Dr. Ruth Arnon
 Institution:  The Weizmann Institute of Science
 Year Elected:  2009
 Class:  2. Biological Sciences
 Subdivision:  209. Neurobiology
 Residency:  International
 Living? :   Living
 Birth Date:  1933
   
 
Prof. Arnon joined the Weizmann Institute of Science in 1960. Prior to her appointment as Vice-President, she served as Head of the Department of Chemical Immunology, and as Dean of the Faculty of Biology. From 1985 to 1994, she was the Director of the Institute's MacArthur Center for Molecular Biology of Tropical Diseases. Prof. Arnon has made significant contributions to the fields of vaccine development, cancer research and to the study of parasitic diseases. Along with Prof. Michael Sela and Dr. Dvora Teitelbaum she developed Copaxone,® a drug for the treatment of multiple sclerosis which was approved by the U.S. Food and Drug Administration, and is presently marketed in the USA, Canada the EU, Australia and many other countries worldwide. Prof. Arnon is a member of the Israel Academy of Sciences, and serves as its Vice-President since 2004. On the world scene, she is an elected member of the European Molecular Biology Organization (EMBO). She has served as President of the European Federation of Immunological Societies (EFIS), and as Secretary-General of the International Union of Immunological Societies (IUIS), as well as in the European Union Research Advisory Board (EURAB). Her awards include the Robert Koch Prize in Medical Sciences, Spain's Jiminez Diaz Memorial Prize, France's Legion of Honor, the Hadassah World Organization's Women of Distinction Award, the Wolf Prize for Medicine, the Rothschild Prize for Biology, and the Israel Prize, Honorary degree – Ben-Gurion University of the Negev, Israel, The AESKU Prize for Life Contribution to Autoimmunity by the 6th International Congress on Autoimmunity, “Yakir” Tel-Hai Academic College, Israel, Elected to the American Philosophical Society. Prof. Arnon is also the Scientific Advisor to the President of the State of Israel. Prof. Arnon is the incumbent of the Paul Ehrlich Chair in Immunochemistry.
 
4Name:  Sir Patrick Bateson
 Institution:  University of Cambridge
 Year Elected:  2006
 Class:  2. Biological Sciences
 Residency:  International
 Living? :   Deceased
 Birth Date:  1938
 Death Date:  August 1, 2017
   
 
Professor Sir Patrick Bateson, FRS, was Emeritus Professor of Ethology, the biological study of behaviour, at the University of Cambridge at the time of his death on August 1, 2017, at the age of 79. He was Provost (Head) of King's College, Cambridge from 1988 to 2003. He was formerly Director of the Sub-Department of Animal Behaviour at Cambridge and later Head of the Department of Zoology. He was Vice-Chairman of the Museums and Galleries Commission and in 2004 was elected President of the Zoological Society of London. He was elected a Fellow of the Royal Society of London in 1983 and was its Biological Secretary and Vice-President from 1998 to 2003. He was knighted in 2003. His research is on the behavioural development of animals, and much of his scientific career has been concerned with bridging the gap between the studies of behaviour and those of underlying mechanisms, focusing on the process of behavioural imprinting in birds. He has also been concerned to link together studies of development and evolution. In addition to his work on birds he has carried out research on behavioural development in mammals, particularly cats, and has supervised field projects on mammals in East Africa. He conducted a research project for the National Trust on the behavioural and physiological effects of hunting red deer with hounds. He has written more than 260 scientific papers and book chapters on behavioural imprinting in birds, the development of play in cats, the development and evolution of behaviour, neural mechanisms of learning, and the conceptual and methodological issues in the study of behaviour and animal welfare. He has also written articles on co-operation, the ethics of using animals in research, and the hunting of red deer with hounds. He has edited 15 books and is co-author (with Paul Martin) of Measuring Behaviour and Design for a Life: How Behaviour Develops.
 
5Name:  Dr. Kamaljit S. Bawa
 Institution:  University of Massachusetts; Ashoka Trust for Research in Ecology and the Environment
 Year Elected:  2019
 Class:  2. Biological Sciences
 Subdivision:  205. Microbiology
 Residency:  International
 Living? :   Living
 Birth Date:  1939
   
 
Kamal Bawa (www.kbawa.com) is Distinguished Professor of Biology at the University of Massachusetts Boston, and Founder-President of the Ashoka Trust for Research in Ecology and the Environment (ATREE). The Ashoka Trust, based in Bangalore, is one of India’s top-ranked environmental think tanks and in 2019 received the UNESCO Sultan Qaboos Prize for Environmental Conservation. He has done extensive work in the Himalaya for a number of years on a wide range of issues from biodiversity conservation to climate change. Kamal Bawa has published more than 200 papers and has authored or edited more than 10 books, and special issues of journals. Among the many awards he has received are: Bullard Fellowship at Harvard University (1972, 2009) Guggenheim Fellowship (1987), Pew Scholar in Conservation and the Environment (1992), Giorgio Ruffolo Fellowship at Harvard University (2009), the Gunnerus Prize in Sustainability Science from the Royal Norwegian Society of Letters and Sciences (2012), the international MIDORI Prize in Biodiversity (2014) from the Aeon Foundation in Japan, the Linnean Medal (2018), and honorary doctorates from the University of Alberta (2014) and Concordia University in Montreal(2019). He is an elected Fellow of the American Academy of Arts and Sciences (2012), the Royal Norwegian Society of Letters and Sciences (2012), and the Royal Society (2015). Kamal Bawa is founding Editor-in Chief of two interdisciplinary journals: Conservation and Society (www.conservationandsociety.org) and Ecology, Economy and Society (http://ecoinsee.org/journal/eb_editors). His latest coffee table book Himalaya: The Mountains of Life, a companion volume to Sahyadri: India’s Western Ghats, was published in 2013.
 
6Name:  Dr. K. Sune D. Bergström
 Institution:  Karolinska Institutet
 Year Elected:  1984
 Class:  2. Biological Sciences
 Subdivision:  201. Molecular Biology and Biochemistry
 Residency:  International
 Living? :   Deceased
 Birth Date:  1916
 Death Date:  August 15, 2004
   
7Name:  Sir Michael J. Berridge
 Institution:  The Babraham Institute; University of Cambridge
 Year Elected:  2007
 Class:  2. Biological Sciences
 Subdivision:  201. Molecular Biology and Biochemistry
 Residency:  International
 Living? :   Deceased
 Birth Date:  1938
 Death Date:  February 13, 2020
   
 
Michael John Berridge was born in 1938 in Gatooma, a small town in the middle of Rhodesia, which is now Zimbabwe. He began his education at Jameson High School where he was fortunate in being taught biology by Pamela Bates who fostered his academic interests and encouraged him to pursue a scientific career. He enrolled in the University of Rhodesia and Nyasaland in Salisbury to read Zoology and Chemistry where he received his B.Sc. (1st Class Honours) in 1960. He then travelled to England to begin research on insect physiology with Sir Vincent Wigglesworth at the University of Cambridge and was awarded his Ph.D. in 1964. He than travelled to the United States to begin a period of post-doctoral study first at the University of Virginia and later at Case Western Reserve University in Cleveland. During his stay in Cleveland he began his interest in how cells communicate with each other and was fortunate to obtain valuable advice from Dr Ted Rall who a few years earlier had worked together with Earl Sutherland who received a Nobel Prize for his discovery of the second messenger cyclic AMP. In 1969 Berridge returned to Cambridge to take up an appointment at the AFRC Unit of Insect Neurophysiology and Pharmacology. He currently is an Emeritus Babraham Fellow at The Babraham Institute Laboratory of Molecular Signalling. Berridge is best known for his discovery of the second messenger inositol trisphosphate (IP3), which plays a universal role in regulating many cellular processes including cell growth and information processing in the nervous system. His studies on cell signalling began with his interest in trying to understand the control of fluid secretion by an insect salivary gland. His introduction and development of this simple model system paved the way for a number of significant observations which culminated in the major breakthrough of uncovering a new second messenger system responsible for regulating intracellular calcium signalling. A role for second messengers in controlling fluid secretion was first recognised when cyclic AMP was found to mimic the stimulatory action of 5-hydroxytryptamine. Subsequent studies revealed that calcium was also important and Berridge was one of the first to draw attention to the integrated action of the cyclic AMP and calcium messenger systems. He showed that signal calcium could be derived from both external and internal reservoirs. A major problem emerged as to how cells gained access to their internal stores of calcium. Berridge provided the first direct evidence to support Michell's hypothesis that the hydrolysis of inositol lipids played a role in calcium signalling. Interest in inositol phosphates began to intensify when Berridge developed a new approach of measuring their formation as a direct way to study receptor-mediated inositol lipid hydrolysis. Of particular significance, was his introduction of the lithium amplification technique to provide an exquisitely sensitive method for measuring inositol lipid turnover. His work on lithium provided new insights into how this drug controls manic-depressive illness. Using the lithium amplification method, Berridge demonstrated that hormones stimulated a rapid formation of IP3, which led him to propose that this metabolite might function as a second messenger. Such a messenger role was rapidly verified when IP3 was found to mobilize calcium when injected into cells. It is now apparent that the IP3/calcium signalling system regulates a wide range of cellular processes such as fertilization, secretion, metabolism, contraction, cell proliferation and information processing in the brain. This work has sparked a worldwide interest in the role of this signalling system in cell regulation. His most recent work has concentrated on the spatial and temporal aspects of calcium signalling. He was one of the first physiologists to provide evidence that the level of calcium might oscillate when cells are stimulated by a hormone. He also showed that oscillation frequency varied with agonist concentration, which led him to propose that the signalling system was frequency-modulated. Berridge's discovery of the IP3/calcium pathway provided an explanation of such oscillatory activity. His laboratory has been at the forefront of recent studies exploiting rapid confocal imaging techniques to characterize the elementary events of calcium signalling. This radically new understanding of how calcium signals are produced has provided new insights into both neural and cardiac cell signalling. Berridge became a Fellow of Trinity College in 1972 and was elected a Fellow of The Royal Society in 1984. In 1999 he was elected to the National Academy of Sciences and the American Academy of Arts and Sciences. For his work on second messengers Berridge has received numerous awards and prizes, including The King Faisal International Prize in Science, The Louis Jeantet Prize in Medicine, The Albert Lasker Medical Research Award, The Heineken Prize for Biochemistry and Biophysics, The Wolf Foundation Prize in Medicine and The Shaw Prize in Life Science and Medicine. In 1998 Berridge was knighted for his service to science.
 
8Name:  Sir Walter Bodmer
 Institution:  Hertford College, Oxford; Weatherall Institute of Molecular Medicine
 Year Elected:  1989
 Class:  2. Biological Sciences
 Subdivision:  204. Medicine, Surgery, Pathology and Immunology
 Residency:  International
 Living? :   Living
 Birth Date:  1936
   
 
Sir Walter Bodmer was principal of Hertford College at Oxford University from 1996-2005. Formerly director general and director of research of the Imperial Cancer Research Fund, he has served as president of the Human Genome Organization (HUGO); president of the British Association for the Advancement of Science; chairman of the BBC Science consultative group; and chairman of the trustees of the Natural History Museum. The co-author of a classic book on human population genetics, he has contributed heavily to both the science and the theory of human genetics. Sir Walter has been a pioneer in elucidating the laws of tissue matches in humans, as well as in the mapping of genes on chromosomes by fusion of human and rodent cells in culture. He led the group that demonstrated that the gene for hereditary colonic polypsis is on chromosome 5. An eminently effective explicator of genetics and a public advocate for science, he has held academic appointments and honorary fellowships at Keble College, Oxford; Clare College, Cambridge; the Royal Colleges of Physicians and of Surgeons; and the Royal Society of Edinburgh. He became a fellow of the Royal Society in 1974, was knighted in 1986 and is a foreign associate of the National Academy of Sciences and a foreign honorary member of the American Academy of Arts & Sciences. He was chancellor of the University of Salford from 1995 to 2005.
 
9Name:  Sir Christopher C. Booth
 Institution:  University College, London
 Year Elected:  1981
 Class:  2. Biological Sciences
 Subdivision:  204. Medicine, Surgery, Pathology and Immunology
 Residency:  International
 Living? :   Deceased
 Birth Date:  1924
 Death Date:  July 13, 2012
   
 
Sir Christopher Booth was a distinguished physician who occupied many high-profile positions, including President of the British Medical Association, Director of the Medical Research Council's Clinical Research Centre and President of the Royal Society of Medicine. His early interests in medical history resulted in a paper on Dr. John Fothergill and angina pectoris that was published in the very first volume of Medical History in 1957 while he was a research registrar at the Hammersmith Hospital, London. Soon after, he commenced important research on vitamin B12 absorption and utilization. In 1971 he published, with Betsy Corner, selected correspondence of John Fothergill, an achievement for any historian, let alone one who was also a professor of medicine. As a historian, Sir Christopher pioneered unique fields, and his enthusiastic investigation of doctors from the Yorkshire Dales revealed a network of remarkable characters, many of them Quakers, whose influence spread from Yorkshire to have great effect in national and international medical worlds. Sir Christopher was an Honorary Professor and member of the Wellcome Trust Group for the study of 20th century medicine at University College, London from 1989 until his death on July 13, 2012. He was elected an international member of the American Philosophical Society in 1981.
 
10Name:  Dr. Sydney Brenner
 Institution:  The Salk Institute
 Year Elected:  1979
 Class:  2. Biological Sciences
 Subdivision:  207. Genetics
 Residency:  International
 Living? :   Deceased
 Birth Date:  1927
 Death Date:  April 5, 2019
   
 
Sydney Brenner was born in South Africa and studied medicine and science at the University of Witwatersrand, Johannesburg. Working in the Physical Chemistry Laboratory, he received his D.Phil. degree from Oxford University in 1952. After briefly returning to South Africa, Dr. Brenner joined the Medical Research Council's Unit in the Cavendish Laboratory at Cambridge. He became the director of its successor, the MRC Laboratory of Molecular Biology, in 1979. In 1987 he became director of the MRC's Unit of Molecular Genetics, retiring in 1992. His early research was in molecular genetics. Working with bacterophages and bacteria, he discovered messenger RNA (working with Jacob and Meselson) and, with Crick, showed that the code was composed of triplets. In the 1960s, he changed the direction of his work and began research on C. elegans, establishing it as a powerful experimental system for the analysis of complex biological processes. Believing that the techniques of cloning and sequencing would open up new ways of approaching genetics, he turned his attention to vertebrate genomics and established the pufferfish genome as a valuable tool in genome analysis. He served as the founder and president of the Molecular Sciences Institute, a private research institute in Berkeley, CA. His achievements were recognized with the Albert Lasker Medical Research Award (1971), both the Royal Medal (1974) and the Copley Medal (1991) from the Royal Society of London, and the Kyoto Prize (1990), to name just four of more than twenty such honors. In 1987 he was named a Companion of Honour. He received the Nobel Prize in Physiology or Medicine in 2002. He was elected to membership in the American Philosophical Society in 1979. Sydney Brenner died April 5, 2019 in Singapore at the age of 92.
 
11Name:  Prof. Erwin Bünning
 Institution:  University of Tübingen
 Year Elected:  1977
 Class:  2. Biological Sciences
 Subdivision:  202. Cellular and Developmental Biology
 Residency:  International
 Living? :   Deceased
 Birth Date:  1906
 Death Date:  10/4/1990
   
12Name:  F. Macfarlane Burnet
 Year Elected:  1960
 Class:  2. Biological Sciences
 Residency:  International
 Living? :   Deceased
 Birth Date:  1899
 Death Date:  8/31/85
   
13Name:  Santiago Ramon y Cajal
 Year Elected:  1932
 Class:  2. Biological Sciences
 Residency:  International
 Living? :   Deceased
 Birth Date:  1852
 Death Date:  10/17/34
   
14Name:  Dr. Torbjörn Caspersson
 Institution:  Karolinska Sjukhuset
 Year Elected:  1974
 Class:  2. Biological Sciences
 Subdivision:  204. Medicine, Surgery, Pathology and Immunology
 Residency:  International
 Living? :   Deceased
 Birth Date:  1910
 Death Date:  December 7, 1997
   
15Name:  Dr. Carlos Chagas
 Institution:  Univeridade Federal de Rio de Janeiro
 Year Elected:  1968
 Class:  2. Biological Sciences
 Subdivision:  209. Neurobiology
 Residency:  International
 Living? :   Deceased
 Birth Date:  1910
 Death Date:  February 16, 2000
   
16Name:  Sir Christopher H. Andrewes
 Year Elected:  1955
 Class:  2. Biological Sciences
 Subdivision:  204. Medicine, Surgery, Pathology and Immunology
 Residency:  International
 Living? :   Deceased
 Birth Date:  1896
 Death Date:  12/31/88
   
17Name:  Dr. Aaron J. Ciechanover
 Institution:  Technion - Israel Institute of Technology
 Year Elected:  2005
 Class:  2. Biological Sciences
 Subdivision:  202. Cellular and Developmental Biology
 Residency:  International
 Living? :   Living
 Birth Date:  1947
   
 
Aaron Ciechanover was born in Haifa, Israel in 1947. He is currently on the academic staff of the Faculty of Medicine of the Technion in Haifa, Israel. He received his M.Sc. (1971) and M.D. (1975) from the Hebrew University in Jerusalem, Israel, and his D.Sc. (1982) from the Technion. There, as a graduate student with Dr. Avram Hershko and in collaboration with Dr. Irwin A. Rose from the Fox Chase Cancer Center in Philadelphia, they discovered that covalent attachment of ubiquitin to the target substrate signals it for degradation. They deciphered the mechanism of conjugation in a cell-free system, described the general proteolytic function of the system in cells, and proposed a model according to which this modification serves as a recognition signal for a specific downstream protease. As a post doctoral fellow with Dr. Harvey Lodish at the Massachusetts Institute of Technology, he collaborated with Drs. Alexander Varshavsky and Daniel Finley, and described the first mutant cell of the system, further corroborating the role of ubiquitin modification as a proteolytic signal in intact cells. Among the many prizes that Dr. Ciechanover received are the 2000 Albert Lasker Award for Basic Medical Research and the 2004 Nobel Prize in Chemistry. Dr. Ciechanover is a member of the Israeli National Academy of Sciences and Humanities, a foreign associate of the National Academy of Sciences of the USA and a member of the Pontifical Academy of Sciences of the Vatican.
 
18Name:  Dr. Bryan C. Clarke
 Institution:  University of Nottingham
 Year Elected:  2003
 Class:  2. Biological Sciences
 Subdivision:  203. Evolution & Ecology, Systematics, Population Genetics, Paleontology, and Physical Anthropology
 Residency:  International
 Living? :   Deceased
 Birth Date:  1932
 Death Date:  February 27, 2014
   
 
Bryan Clarke had a distinguished career as a leading figure in the genetical study of populations in nature. More than any other he demonstrated the widespread occurence and importance of frequency-dependent selection. This work, both theoretical and empirical and strikingly original, has helped to explain such diverse phenomena as clinal variation in gene frequencies, evolutionary dynamics of parasite-host interactions, speciation mechanisms in snails, and the maintenance of genetic variation in human populations. He gained prominence as an international spokesman for ecological genetics and as advisor to organizations including the International Union for the Conservation of Nature. Dr. Clarke served on the faculty of the University of Nottingham since 1971 and became Professor of Genetics Emeritus in 1997. He was awarded the Linnean Society of London's Darwin-Wallace Medal in 2008 and the Royal Society's Darwin Medal in 2010. Bryan Clarke died February 27, 2014, at the age of 81 in Nottingham, UK.
 
19Name:  Prof. Francis H. C. Crick
 Institution:  Salk Institute & University of California, San Diego
 Year Elected:  1972
 Class:  2. Biological Sciences
 Subdivision:  209. Neurobiology
 Residency:  International
 Living? :   Deceased
 Birth Date:  1916
 Death Date:  July 28, 2004
   
20Name:  Henry H. Dale
 Year Elected:  1939
 Class:  2. Biological Sciences
 Residency:  International
 Living? :   Deceased
 Birth Date:  1875
 Death Date:  7/23/68
   
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