American Philosophical Society
Member History

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Class
2. Biological Sciences[X]
281Name:  Archibald V. Hill
 Year Elected:  1938
 Class:  2. Biological Sciences
 Residency:  International
 Living? :   Deceased
 Birth Date:  1886
 Death Date:  6/3/77
   
282Name:  Dr. Maurice R. Hilleman
 Institution:  Merck Institute for Vaccinology & University of Pennsylvania
 Year Elected:  1997
 Class:  2. Biological Sciences
 Subdivision:  205. Microbiology
 Residency:  Resident
 Living? :   Deceased
 Birth Date:  1919
 Death Date:  April 11, 2005
   
 
Dr. Maurice R. Hilleman is Director, Merck Institute for Vaccinology and Adjunct Professor of Pediatrics, University of Pennsylvania. His past career of six decades of medical research included the Squibb Research Labs, Walter Reed Army Institute of Research, the Rockefeller Institute for Medical Research, and the Merck Institute for Medical Research. His bibliography includes more than 500 original publications in virology, immunology, epidemiology, and infectious diseases. Dr. Hilleman is a senior statesman and authority in the medical sciences for basic discoveries and vaccine developments. He has received numerous awards and accolades from academia, government, and industry. Among the most significant, he is an elected member of the U.S. National Academy of Sciences; the Institute of Medicine of the Academy; the American Philosophical Society, and the American Academy of Arts and Sciences. Distinctive honors include the Lasker Medical Research Award; Award of the National Medal of Science by President Reagan; the Robert Koch Gold Medal (Berlin); the Prince Mahidol Award presented by the King of Thailand; the Maxwell Finland Award; the Albert B. Sabin Gold Medal; Decoration for Distinguished Science Achievement by the U.S. Secretary of Defense, and numerous lifetime achievement awards. Dr. Hilleman received his B.S. degree from Montana State University (1941), and a Ph.D. from the University of Chicago (1944). He holds honorary doctorate degrees from U.S. and foreign universities. Dr. Hilleman’s career has been devoted to both basic and applied research with breakthrough discoveries and developments in virology, cancer, immunology, epidemiology, and vaccinology. Basic research examples include the discoveries of SV40 virus and its oncogenicity, the codiscoveries of the Adenoviruses and the Rhinoviruses, purification and characterization of interferon and it’s induction by double-stranded RNA, pioneering propagation of hepatitis A virus and its growth in cell culture. He pioneered the development of numerous live, killed and recombinant vaccines including measles, mumps, rubella, MMR, varicella, Marek’s Disease, hepatitis A; both plasma-derived and recombinant hepatitis B, and the commercial evolution of vaccines against meningococci and pneumococci. He has been credited with developing more vaccines than any person and is recognized for having changed the face of the world in providing means to prevent and control a number of its most important diseases. Many consider him a living legend!
 
283Name:  Sir Harold Himsworth
 Institution:  Medical Research Council
 Year Elected:  1972
 Class:  2. Biological Sciences
 Subdivision:  204. Medicine, Surgery, Pathology and Immunology
 Residency:  International
 Living? :   Deceased
 Birth Date:  1905
 Death Date:  11/1/93
   
284Name:  Dr. Peter H. von Hippel
 Institution:  Institute of Molecular Biology, University of Oregon
 Year Elected:  2004
 Class:  2. Biological Sciences
 Subdivision:  206. Physiology, Biophysics, and Pharmacology
 Residency:  Resident
 Living? :   Living
 Birth Date:  1931
   
 
Peter von Hippel was born in Germany and became a naturalized US citizen in 1942. He obtained his BS, MS, and Ph. D. degrees at MIT, working on the physical biochemistry of protein complexes in the laboratory of Professor David F. Waugh. He then did postdoctoral work on actomyosin complexes with Dr. Manuel F. Morales at the Naval Medical Research Institute in Bethesda, followed by a period as a staff scientist at NMRI while serving in the U.S. Navy He began his academic career in 1959 as an Assistant Professor of Biochemistry at Dartmouth Medical School in Hanover, New Hampshire. His work at that time involved analyzes of the structure of proteins and nucleic acids, and the effects on these macromolecules of ions and other solvent additives. He remained at Dartmouth until 1967, and then moved to the University of Oregon as a Professor of Chemistry and Member of the Institute of Molecular Biology, where he has been ever since. While at Oregon the research program in the von Hippel laboratory has progressed from studies of the interactions of simple regulatory proteins and protein models with DNA to the quantitative analysis of the structure and function of various macromolecular complexes involved in the control of DNA replication and RNA transcription. He has served as Director of the Institute of Molecular Biology and Chair of the Chemistry Department at Oregon, and in 1989 was appointed an American Cancer Society Research Professor of Chemistry, which has spared him from further formal administrative activities. In other venues Dr. von Hippel has served on various study sections and advisory committees for both NIH and NSF and has participated in the activities of various professional organizations, including serving on the Board of Directors of FASEB and as President of the Biophysical Society. He has served on the editorial boards of numerous professional journals, and he and his laboratory colleagues have published more than 240 research papers. Dr. von Hippel was elected to membership in the National Academy of Sciences in 1978, to fellowship of the American Academy of Arts & Sciences in 1979, and to resident membership in the American Philosophical Society in 2004. He is the 2021 recipient of the Ignacio Tinoco Award from the Biophysical Society for his exceptional contributions to the field of biophysics.
 
285Name:  Frederick L. Hisaw
 Year Elected:  1940
 Class:  2. Biological Sciences
 Residency:  Resident
 Living? :   Deceased
 Birth Date:  1891
 Death Date:  12/03/72
   
286Name:  William H. Hobbs
 Year Elected:  1909
 Class:  2. Biological Sciences
 Residency:  Resident
 Living? :   Deceased
 Birth Date:  1865
 Death Date:  01/01/53
   
287Name:  Sir Alan L. Hodgkin
 Year Elected:  1967
 Class:  2. Biological Sciences
 Subdivision:  206. Physiology, Biophysics, and Pharmacology
 Residency:  International
 Living? :   Deceased
 Birth Date:  1914
 Death Date:  12/20/98
   
288Name:  Dr. Hopi E. Hoekstra
 Institution:  Harvard University; Howard Hughes Medical Institute
 Year Elected:  2018
 Class:  2. Biological Sciences
 Subdivision:  203. Evolution & Ecology, Systematics, Population Genetics, Paleontology, and Physical Anthropology
 Residency:  Resident
 Living? :   Living
 Birth Date:  1972
   
 
Hopi E. Hoekstra is the Alexander Agassiz Professor of Zoology in the Departments of Organismic & Evolutionary Biology and the Molecular & Cellular Biology at Harvard University. She is the Curator of Mammals in the Museum of Comparative Zoology, an Institute Member at the Broad Institute and an Investigator at the Howard Hughes Medical Institute. Her research focuses on understanding the evolution and genetics of morphological and behavioral traits that affect fitness of individuals in the wild. Using deer mice as a model system, she first dissected the molecular, genetic and developmental basis of camouflaging coloration to understand the mechanisms driving adaptation. Later, she focused on unraveling the genetic and neurobiological underpinnings of complex natural behaviors. She received her B.A. from UC Berkeley and her Ph.D. from the University of Washington. She has received Young Investigator awards from the American Society of Naturalists and the Arnold and Mabel Beckman Foundation, and most recently, the Lounsbery Medal from the National Academy of Sciences (2015). She gave the 2013 Commencement speech at UC Berkeley’s Integrative Biology Department and has been profiled in The New York Times. In 2016, she was elected into the National Academy of Sciences and in 2017, the American Academy of Arts & Sciences. She also teaches in Harvard’s introductory Life Science course Genetics, Genomics and Evolution to approximately 500 freshmen each year, and has been awarded the Fannie Cox Prize and a Harvard College Professorship for teaching excellence.
 
289Name:  Dr. Bert Hölldobler
 Institution:  Biozentrum of the University of Würzburg; Arizona State University
 Year Elected:  1997
 Class:  2. Biological Sciences
 Subdivision:  205. Microbiology
 Residency:  International
 Living? :   Living
 Birth Date:  1936
   
 
Bert Hölldobler is one of the foremost authorities in the world on insect behavior and the behavioral ecology of invertebrate animals. In a series of brilliant experiments and field studies over the past forty years, he has demonstrated the extreme adaptations to the environment of which the insect brain is capable: for example, the demonstration of military-like strategies and "diplomacy" in conflict between ant colonies and the basis of ant trap jaw predation -- the reflex arc and most rapid mechanical movements known in animals. His Pulitzer Prize-winning book (with E.O. Wilson) The Ants is a widely-hailed classic, and The Superorganism, Wilson's and Hölldobler's most recent collaboration, provides another in-depth look at the intricate ways of social insects. As Professor of Zoology Emeritus at the Biozentrum of the University of Würzburg and Foundation Professor of Life Sciences at the Arizona State University School of Life Sciences, Dr. Hölldobler continues his interest in the evolution of social organizations in insects and in the underlying mechanisms that make insect societies work. In 2016 he was awarded the Lorenz Oken Medal.
 
290Name:  Dr. Leroy Hood
 Institution:  Phenome Health; Institute for Systems Biology
 Year Elected:  2000
 Class:  2. Biological Sciences
 Subdivision:  201. Molecular Biology and Biochemistry
 Residency:  Resident
 Living? :   Living
 Birth Date:  1938
   
 
Leroy E. Hood is the President and Director of the Institute for Systems Biology, a not-for-profit institution he recently established. He has helped start more than half a dozen companies, including Amgen, the largest biotech company, and Applied Biosystems, the leading maker of genetic analysis equipment. He received an M.D. at the Johns Hopkins School of Medicine in 1964 and a Ph.D. in biochemistry at California Institute of Technology in 1968. A member of the faculty of the California Institute of Technology for twenty-two years, he was also director of the Cancer Center, 1981-90, and director of the NSF Science and Technology Center for Molecular Biotechnolgy, 1989-92. From 1992-2000 he was the Director of the NSF Science and Technology Center for Molecular Biotechnology at the University of Washington, as well as William Gates III Professor and Chairman of the Department of Molecular Biotechnology, professor in the Departments of Bioengineering and Immunology, and an adjunct professor in the Departments of Medicine and Computer Science. Leroy Hood played a central role in deciphering the mechanisms of immunological diversity by being among the first to clone and characterize genes encoding antibodies, genes of the major histocompatibility complex, and T-cell receptors. His laboratory also developed four instruments widely used to synthesize and sequence genes and proteins. Dr. Hood also played a pioneering role in the Human Genome Project and co-edited The Code of Codes, covering the scientific, legal, and ethical aspects of the Human Genome Project. Dr. Hood initiated major programs for bringing hands-on, inquiry-based science to all levels of teachers in Seattle. Dr. Hood is the recipient of many awards, including the Louis Pasteur Award, Dickson Prize, Lasker Award, Rabbi Shai Shacknai Memorial Prize of Hebrew University, the American College of Physicians Award, the NAE's Fritz J. and Dolores H. Russ Prize, and the National Medal of Science (2012). In January 2017 he was awarded the National Academy of Sciences' Award for Chemistry in Service to Society. He is a member of the National Academy of Sciences and American Academy of Arts & Sciences and was elected a member of the American Philosophical Society in 2000.
 
291Name:  Sir Frederick G. Hopkins
 Year Elected:  1937
 Class:  2. Biological Sciences
 Residency:  International
 Living? :   Deceased
 Birth Date:  1861
 Death Date:  5/16/47
   
292Name:  Frank Lappin Horsfall
 Year Elected:  1956
 Class:  2. Biological Sciences
 Residency:  Resident
 Living? :   Deceased
 Birth Date:  1906
 Death Date:  2/19/71
   
293Name:  Dr. H. Robert Horvitz
 Institution:  Massachusetts Institute of Technology; Howard Hughes Medical Institute; McGovern Institute for Brain Research
 Year Elected:  2004
 Class:  2. Biological Sciences
 Subdivision:  207. Genetics
 Residency:  Resident
 Living? :   Living
 Birth Date:  1947
   
 
H. Robert Horvitz's Nobel Prize in Physiology and Medicine, awarded for his pioneering genetic dissection of programmed cell death (apoptosis), including the crucial discovery of the first caspase that mediates apoptosis, celebrated just one of his several comparably important contributions. Through genetic analysis of the nematode Caenorhabditis elegans, Dr. Horvitz discovered and dissected many genes and pathways that play highly specific roles during animal development, and in animal behavior as well. He defined genes that control specific aspects of cell lineage and cell fate, including the generation of cell diversity during development; the timing of particular developmental events; inter- and intracellular signaling; and programmed cell death. Dr. Horvitz's molecular analyses of these genes revealed most of them to be strikingly similar to genes found in other organisms, including humans, and in many cases similar to genes that cause human disease. A member of the Massachusetts Institute of Technology faculty since 1978, Dr. Horvitz has been David H. Koch Professor of Cancer Biology since 2001. He holds a Ph.D. from Harvard University (1974) and is a member of the National Academy of Sciences (1991); the American Academy of Arts & Sciences (1994); and the Genetics Society of America (president, 1995).
 
294Name:  Dr. Susan Band Horwitz
 Institution:  Albert Einstein College of Medicine
 Year Elected:  2013
 Class:  2. Biological Sciences
 Subdivision:  201. Molecular Biology and Biochemistry
 Residency:  Resident
 Living? :   Living
 Birth Date:  1937
   
 
Dr. Susan Band Horwitz was born in Boston where she grew up and attended public high school. After graduating from Bryn Mawr College, she attended Brandeis University where she received her Ph.D. in Biochemistry. She was a postdoctoral fellow in the Departments of Pharmacology at Tufts University Medical School, Emory University Medical School and the Albert Einstein College of Medicine. She joined the faculty at Albert Einstein in 1970 and is presently the Falkenstein Professor of Cancer Research, Co-Chair of the Department of Molecular Pharmacology and the Associate Director for Therapeutics at the Albert Einstein Cancer Center. Dr. Horwitz has had a continuing interest in natural products as a source of new drugs for the treatment of cancer. Her contributions span several decades of research and encompass agents which have served as prototypes for some of our most important drugs that are currently in clinical use. She made major contributions to our understanding of the mechanisms of action of camptothecin, the epipodophyllotoxins and bleomycin. However, Dr. Horwitz’ most seminal research contribution has been in the development of Taxol, a drug isolated from the yew plant, Taxus brevifolia. Although no one was interested in Taxol when she began her studies, today it is an important antitumor drug approved by the FDA for the treatment of ovarian, breast and lung carcinomas. The drug has been given to over a million patients. Dr. Horwitz' research played an important role in encouraging the development of Taxol by the National Cancer Institute. Dr. Horwitz is a member of the National Academy of Sciences, the Institute of Medicine, and the American Academy of Arts and Sciences. She has received numerous honors and awards including the Cain Memorial Award of the AACR, the ASPET Award for Experimental Therapeutics, the C. Chester Stock Award from Memorial Sloan-Kettering Cancer Center, the Warren Alpert Foundation Prize from Harvard Medical School, The American Cancer Society’s Medal of Honor -Clinical Research Award, The Mayor's Award for Excellence in Science & Technology, the Barnard Medal of Distinction, and the 2014 John Scott Science Award. In 2011, Dr. Horwitz received the AACR Lifetime Achievement Award in Cancer Research and The New York Academy Medal for Distinguished Contributions in Biomedical Science. She served as president (2002-2003) of the American Association of Cancer Research. Susan Horwitz was elected a member of the American Philosophical Society in 2013.
 
295Name:  Carl I. Hovland
 Year Elected:  1950
 Class:  2. Biological Sciences
 Residency:  Resident
 Living? :   Deceased
 Birth Date:  1913
 Death Date:  04/16/61
   
296Name:  Leland O. Howard
 Year Elected:  1911
 Class:  2. Biological Sciences
 Residency:  Resident
 Living? :   Deceased
 Birth Date:  1858
 Death Date:  05/01/50
   
297Name:  William H. Howell
 Year Elected:  1903
 Class:  2. Biological Sciences
 Residency:  Resident
 Living? :   Deceased
 Birth Date:  1860
 Death Date:  02/06/45
   
298Name:  Dr. F. Clark Howell
 Institution:  University of California, Berkeley
 Year Elected:  1975
 Class:  2. Biological Sciences
 Subdivision:  203. Evolution & Ecology, Systematics, Population Genetics, Paleontology, and Physical Anthropology
 Residency:  Resident
 Living? :   Deceased
 Birth Date:  1925
 Death Date:  March 10, 2007
   
299Name:  Ales Hrdlicka
 Year Elected:  1918
 Class:  2. Biological Sciences
 Residency:  Resident
 Living? :   Deceased
 Birth Date:  1869
 Death Date:  09/05/43
   
300Name:  Dr. Sarah Blaffer Hrdy
 Institution:  University of California, Davis
 Year Elected:  2011
 Class:  2. Biological Sciences
 Subdivision:  205. Microbiology
 Residency:  Resident
 Living? :   Living
 Birth Date:  1946
   
 
Sarah Blaffer Hrdy graduated summa cum laude from Radcliffe College and earned her PhD at Harvard in 1975. Currently she is professor emerita in the Department of Anthropology at the University of California, Davis. She has been elected to the California Academy of Sciences, the National Academy of Sciences and the American Academy of Arts and Sciences. She is the sole author of five books including The Woman That Never Evolved (1981), Mother Nature (1999), and Mothers and Others: The evolutionary origins of mutual understanding (2009) as well as co-editor of Infanticide: Comparative and Evolutionary Perspectives and Attachment and Bonding: A New Synthesis. Her current focus is on how evolutionary perspectives can help us better understand the needs of children. She and her husband, a medical doctor, have three children and currently combine growing walnuts with habitat restoration on their farm in northern California (which can be found at www.citrona.com). In 2014 she was received the NAS Award for Scientific Publishing.
 
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