APS Member History

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201	Name:	Clarence Henry Graham
	Year Elected:	1956
	Class:	2. Biological Sciences
	Residency:	Resident
	Living? :	Deceased
	Birth Date:	1906
	Death Date:	7/25/1971

202	Name:	Dr. Peter R. Grant
	Institution:	Princeton University
	Year Elected:	1991
	Class:	2. Biological Sciences
	Subdivision:	203. Evolution & Ecology, Systematics, Population Genetics, Paleontology, and Physical Anthropology
	Residency:	Resident
	Living? :	Living
	Birth Date:	1936

	A key figure in modern evolutionary biology, Peter R. Grant is most interested in ecology, evolution and behavior. He specializes in the finches of the Galapagos Islands that were originally studied by Charles Darwin and has conducted long-term studies of individual finch populations on some of the smaller islands. He has been able to show that, even over a space of time as short as 15 years, there have been numerous climate changes which have led to clear evidence of natural selection influencing and modifying the genetically determined physical characteristics of the indiviudal finches within a population. With numerous publications to his credit, Dr. Grant has been a professor of ecology and evolutionary biology at Princeton University since 1985. He was educated at the University of Cambridge, the University of British Columbia and Yale University and previously taught at McGill University and the University of Michigan. He has received awards including a Guggenheim Fellowship (1985-86), an Alexander von Humboldt Fellowship (1996) and the American Ornithologists' Union's Brewster Medal (1983). He is a fellow of the Royal Society of London (1987), the Royal Society of Canada and the American Academy of Arts & Sciences (1997) and a foreign associate of the National Academy of Sciences (2007). He has also shared numerous awards with his wife, fellow evolutionary ecologist Rosemary Grant. These include the Academy of Natural Science's Leidy Medal (1994), the E.O. Wilson Prize of the American Society of Naturalists (1998), the Darwin Medal for Evolutionary Biology (2003), the A.I.B.S. Outstanding Scientist Award (2005), the Balzan Prize in Population Biology (2005), the Linnean Society of London's Darwin-Wallace Medal (2008), the Kyoto Prize from the Inamori Foundation (2009), the Royal Medal in Biology from the Royal Society of London (2017), and the BBVA Foundation Frontiers of Knowledge Award (2017)

203	Name:	Dr. B. Rosemary Grant
	Institution:	Princeton University
	Year Elected:	2010
	Class:	2. Biological Sciences
	Subdivision:	203. Evolution & Ecology, Systematics, Population Genetics, Paleontology, and Physical Anthropology
	Residency:	Resident
	Living? :	Living
	Birth Date:	1936

	Rosemary Grant is interested in the Darwinian problem of explaining how new species arise. In 1973 she and her husband Peter began a long-term and continuing study of the adaptive radiation of Darwin’s finches on the Galápagos Islands. Her research combines studies of archipelago-wide patterns of evolution with detailed investigations of ecological, behavioral and genetic mechanisms of change on the two islands of Genovesa and Daphne. This work has been published in numerous scientific papers and two books, the most recent being How and Why Species Multiply (Princeton University Press 2008). Rosemary Grant was initially trained at the University of Edinburgh, received a PhD from Uppsala University in Sweden, and was Senior Research Scholar with rank of Professor in the Department of Ecology and Evolution at Princeton University. She is now emeritus. She has also shared numerous awards with her husband, fellow evolutionary ecologist Peter Grant. These include the Academy of Natural Science's Leidy Medal (1994), the E.O. Wilson Prize of the American Society of Naturalists (1998), the Darwin Medal for Evolutionary Biology (2003), the A.I.B.S. Outstanding Scientist Award (2005), the Balzan Prize in Population Biology (2005), the Linnean Society of London's Darwin-Wallace Medal (2008), the Kyoto Prize from the Inamori Foundation (2009), the Royal Medal in Biology from the Royal Society of London (2017), and the BBVA Foundation Frontiers of Knowledge Award (2017).

204	Name:	Dr. Ann M. Graybiel
	Institution:	Massachusetts Institute of Technology
	Year Elected:	2016
	Class:	2. Biological Sciences
	Subdivision:	208. Plant Sciences
	Residency:	Resident
	Living? :	Living
	Birth Date:	1942

	Ann Martin Graybiel and coworkers pioneered understanding of the basal ganglia, brain structures related to movement and emotion that are disordered in neurologic and neuropsychiatric disorders. Relatively little was known about the neurobiology of these structures until Graybiel and her students discovered the structural architecture of the striatum, a physical design now known to underpin the organization of genes and neurotransmitters, including dopamine, linked to Parkinson’s disease. By training animals to learn habits, she and her group discovered neural activity templates for habit learning in the striatum and found that distinct activity patterns uniquely characterize different motor and emotion-related regions. Graybiel and students now are finding that these templates can be modified by circuit intervention, opening the possibility of new therapeutic approaches to disorders of movement and emotion.

205	Name:	Dr. Paul Greengard
	Institution:	Rockefeller University
	Year Elected:	1994
	Class:	2. Biological Sciences
	Subdivision:	208. Plant Sciences
	Residency:	Resident
	Living? :	Deceased
	Birth Date:	1925
	Death Date:	April 13, 2019

	Paul Greengard was Vincent Astor Professor and Head of the Laboratory of Molecular and Cellular Neuroscience at The Rockefeller University from 1983 until his death in 2019. He received his Ph.D. degree from Johns Hopkins University in 1953. He then spent five years in England receiving advanced training at the University of London, at Cambridge University and at the National Institute of Medical Research. Upon his return to the United States, Greengard worked as Director of the Department of Biochemistry at Geigy Research Laboratories, in Ardsley, New York, for eight years. From 1968 to 1983, he served as Professor of Pharmacology and Psychiatry at Yale University. Greengard authored over 1,000 scientific publications and his achievements earned him numerous prestigious awards. He was awarded the Nobel Prize in Physiology or Medicine in 2000 for his discovery of how dopamine and a number of other neurotransmitters exert their action in the brain. Paul Greengard died April 13, 2019 in Manhattan at the age of 93.

206	Name:	Alan Gregg
	Year Elected:	1944
	Class:	2. Biological Sciences
	Residency:	Resident
	Living? :	Deceased
	Birth Date:	1890
	Death Date:	6/19/57

207	Name:	Herbert E. Gregory
	Year Elected:	1923
	Class:	2. Biological Sciences
	Residency:	Resident
	Living? :	Deceased
	Birth Date:	1870

208	Name:	William K. Gregory
	Year Elected:	1925
	Class:	2. Biological Sciences
	Residency:	Resident
	Living? :	Deceased
	Birth Date:	1876

209	Name:	Dr. Carol W. Greider
	Institution:	University of California Santa Cruz
	Year Elected:	2016
	Class:	2. Biological Sciences
	Subdivision:	201. Molecular Biology and Biochemistry
	Residency:	Resident
	Living? :	Living
	Birth Date:	1961

	Carol Greider, Ph.D. received her bachelor’s degree from the University of California at Santa Barbara in 1983 and a Ph.D. in 1987 from the University of California at Berkeley. In 1984, working together with Dr. Elizabeth Blackburn, she discovered telomerase, an enzyme that maintains telomeres, or chromosome ends. In 1988, Dr. Greider was recruited to Cold Spring Harbor Laboratory as an independent Cold Spring Harbor Fellow, where she cloned and characterized the RNA component of telomerase. In 1990, Dr. Greider was appointed as an assistant investigator at Cold Spring Harbor Laboratory, followed later by appointment to Investigator in 1994. She expanded the focus of her telomere research to include the role of short telomeres in cellular senescence, cell death and in cancer. In 1997, Dr. Greider moved her laboratory to the Department of Molecular Biology and Genetics at Johns Hopkins University School of Medicine. In 2003, she was appointed as the Daniel Nathans Professor and Director of the Department of Molecular Biology and Genetics. Dr. Greider’s group continued to study the biochemistry of telomerase and determined the secondary structure of the human telomerase RNA. In addition, she characterized the loss of telomere function in mice, which allowed an understanding of human diseases that make up the short telomere syndromes. Dr. Greider shared the Nobel Prize in Physiology or Medicine in 2009 with Drs. Elizabeth Blackburn and Jack Szostak for their work on telomeres and telomerase. In 2014, Dr. Greider was appointed as a Blooomberg Distinguished Professor at Johns Hopkins University. Dr. Greider directs a group of scientists studying both the role of short telomeres in age-related disease and cancer as well as the regulatory mechanism that maintain telomere length. In 2020 she became Distinguished Professor of Molecular, Cell, and Developmental (MCD) Biology at University of California Santa Cruz.

210	Name:	Dr. Donald R. Griffin
	Institution:	Rockefeller University
	Year Elected:	1971
	Class:	2. Biological Sciences
	Subdivision:	210. Behavioral Biology, Psychology, Ethology, and Animal Behavior
	Residency:	Resident
	Living? :	Deceased
	Birth Date:	1915
	Death Date:	November 7, 2003

211	Name:	Dr. Diane E. Griffin
	Institution:	Johns Hopkins University
	Year Elected:	2018
	Class:	2. Biological Sciences
	Subdivision:	205. Microbiology
	Residency:	Resident
	Living? :	Living
	Birth Date:	1940

	Diane E. Griffin MD, PhD is University Distinguished Service Professor and former Chair of the W. Harry Feinstone Department of Molecular Microbiology and Immunology at the Johns Hopkins Bloomberg School of Public Health and Vice President of the US National Academy of Sciences. She earned her BA in Biology at Augustana College in Rock Island, IL and her MD and PhD at Stanford University School of Medicine. Her research interests are in the area of pathogenesis of viral diseases with a particular focus on measles and arboviral encephalitis. These studies address issues related to virulence and the role of immune responses in protection from infection and in clearance of infection. She has more than 400 publications and has served on multiple advisory and editorial boards. She is the US Chair of the US-Japan Cooperative Medical Sciences Program and past president of the American Society for Virology and the American Society for Microbiology (ASM). She is a member of the National Academy of Medicine and Association of American Physicians, as well as the National Academy of Sciences. Among other honors, she has received the Rudolf Virchow Medal from the University of Wurzburg (2010), Wallace Sterling Lifetime Alumni Achievement Award from Stanford University (2011), FASEB Excellence in Science Award (2015), Maxwell Finland Award from the NFID (2016) and MilliporeSigma Alice C. Evans Award from the ASM (2017).

212	Name:	J. P. Crozer Griffith
	Year Elected:	1907
	Class:	2. Biological Sciences
	Residency:	Resident
	Living? :	Deceased
	Birth Date:	1856

213	Name:	Dr. David A. Hamburg
	Institution:	Weill Cornell Medical College; Carnegie Corporation of New York
	Year Elected:	1983
	Class:	2. Biological Sciences
	Subdivision:	205. Microbiology
	Residency:	Resident
	Living? :	Deceased
	Birth Date:	1925
	Death Date:	April 21, 2019

	David Hamburg was president emeritus at Carnegie Corporation of New York, where he served as the Corporation's eleventh president from 1982-97. Under his leadership the work of the Corporation focused on education and healthy development of children and youth, human resources in developing countries, and international security issues. He established a number of task forces on education and preventing conflict which produced seminal research and policy analysis and which will continue to influence the work in these fields in the future. A medical doctor, Dr. Hamburg had a long history of leadership in the research, medical and psychiatric fields before his transition from a trustee of Carnegie to its president. An authority on psychosomatic and psychiatric diseases, he was broadly interested in human genetics and evolution. He was chief of the adult psychiatry branch at the National Institutes of Health, from 1958-61; professor and chairman of the department of psychiatry and behavioral sciences at Stanford University from 1961-72; Reed-Hodgson Professor of Human Biology at Stanford University from 1972-76; president of the Institute of Medicine, National Academy of Sciences, 1975-80; and director of the division of health policy research and education and John D. MacArthur Professor of Health Policy at Harvard University, 1980-83. He served as president and chairman of the board (1984-1986) of the American Association for the Advancement of Science. Hamburg was a member of the United States Defense Policy Board with Secretary of Defense William Perry and cochair with former Secretary of State Cyrus Vance of the Carnegie Commission on Preventing Deadly Conflict. He was a member of President Clinton's Committee of Advisors on Science and Technology and a visiting professor at Harvard Medical School's department of social medicine and was the founder of the Carnegie Commission on Science, Technology and Government. In May 2006 Secretary-General Kofi Annan appointed him to chair the newly formed United Nations Advisory Committee on Genocide Prevention. The committee provided guidance and support to the work of the UN Special Advisor on the Prevention of Genocide and contributed to the broader efforts of the UN to avert massive crimes against humanity. He was DeWitt Wallace Distinguished Scholar at the Weill Cornell Medical College and Co-Chair of the Social Medicine and Public Policy Programs. Hamburg received both his A.B. and M.D. degrees from Indiana University. He also received numerous honorary degrees during his career as well as the American Psychiatric Association's Distinguished Service Award in 1991, the Presidential Medal of Freedom at the White House in 1996, the International Peace Academy's 25th Anniversary Special Award in 1996, the Achievement in Children and Public Policy Award from the Society for Research in Child Development in 1997, and the National Academy of Sciences' Public Welfare Medal in 1998. In 2007 he received the Institute of Medicine's Rhoda and Bernard Sarnat International Award in Mental Health jointly with his wife Beatrix; similarly, they were jointly awarded the 2015 Pardes Humanitarian Prize in Mental Health from the Brain & Behavior Research Foundation. David Hamburg died on April 21, 2019 in Washington, D.C. at the age of 93.

214	Name:	Dr. Garrett Hardin
	Institution:	University of California, Santa Barbara
	Year Elected:	1974
	Class:	2. Biological Sciences
	Subdivision:	203. Evolution & Ecology, Systematics, Population Genetics, Paleontology, and Physical Anthropology
	Residency:	Resident
	Living? :	Deceased
	Birth Date:	1915
	Death Date:	September 14, 2003

215	Name:	Harry F. Harlow
	Year Elected:	1957
	Class:	2. Biological Sciences
	Residency:	Resident
	Living? :	Deceased
	Birth Date:	1905
	Death Date:	12/6/81

216	Name:	Robert A. Harper
	Year Elected:	1909
	Class:	2. Biological Sciences
	Residency:	Resident
	Living? :	Deceased
	Birth Date:	1862
	Death Date:	05/12/46

217	Name:	J. George Harrar
	Year Elected:	1962
	Class:	2. Biological Sciences
	Residency:	Resident
	Living? :	Deceased
	Birth Date:	1906
	Death Date:	04/18/82

218	Name:	Ross G. Harrison
	Year Elected:	1913
	Class:	2. Biological Sciences
	Residency:	Resident
	Living? :	Deceased
	Birth Date:	1870
	Death Date:	09/30/59

219	Name:	Dr. Stephen Coplan Harrison
	Institution:	Harvard University & Howard Hughes Medical Institute
	Year Elected:	1997
	Class:	2. Biological Sciences
	Subdivision:	206. Physiology, Biophysics, and Pharmacology
	Residency:	Resident
	Living? :	Living
	Birth Date:	1943

	Stephen Harrison is a world leader in understanding virus structure and in probing the relationship between structure and function of complex protein assemblies. He has devised innovative methodological enhancements of X-ray crystallography to determine the detailed molecular structures of viruses, cell-surface receptors, and DNA-protein complexes. His pioneering studies of small plant viruses at atomic dimensions revealed the basic molecular design of a large class of RNA viruses of plants, insects, and vertebrates. This work, and his subsequent studies of many other viruses, allowed Dr. Harrison to formulate principles that govern viral structure, assembly, stability, cellular attachment, and fusion. This information is fundamental to understanding viral disease and to the design of antiviral drugs and vaccines. Similarly, Dr. Harrison's X-ray crystallographic studies of the structures of DNA-protein complexes revealed important molecular mechanisms in the control of gene activity. Dr. Harrison earned his Ph.D. from Harvard University in 1967. He joined the Harvard faculty in 1971 and is currently director of the Center for Molecular and Cellular Dynamics at Harvard Medical School and head of the Laboratory of Molecular Medicine at Children's Hospital, Boston. He won the Welch Award in Chemistry in 2015 and the Rosenstiel Award for Basic Medical Research in 2018.

220	Name:	Haldan Keffer Hartline
	Year Elected:	1952
	Class:	2. Biological Sciences
	Residency:	Resident
	Living? :	Deceased
	Birth Date:	1903
	Death Date:	3/17/83

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